New Five Southern Tethyan Agglutinated Foraminiferal Species

Haidar Salim Anan

Al Azhar University-Gaza, P.O Box 1277, Gaza, Palestine


Received: 27 June 2022; Revised: 13 September 2022; Accepted: 3 January 2023; Published Online: 16 February 2023

Copyright © 2022 Author(s). Published by Nan Yang Academy of Sciences Pte. Ltd.

Creative Commons LicenseThis is an open access article under the Creative Commons Attribution-NonCommercial 4.0 International (CC BY-NC 4.0) License.


Benthic foraminifera was studied as part of an investigation of the Dababiya section (Egypt), which was undertaken to document a candidate section for the Global Stratotype Section and Point (GSSP) for the base of the Ypresian Stage, Thanetian/Ypresian boundary, Late Campanian Dokan and Azmer sections of Kurdistan (NE Iraq), and the type section of Khangiran Formation of Khangiran section (NE Iran). Five new species of the genera Pseudogaudryina and Pseudogaudryinella are proposed, from three countries Egypt, Iraq and Iran, and from Campanian and Ypresian sediments. Three species of Pseudogaudryina: P. dababiyaensis (from Egypt), P. kurdistanensis and P. iraqensis (from Iraq), two species from Iran: Pseudogaudryinella iranica and Pseudoclavulina iranica. The Tethyan continental shelf fauna Midway-Type Fauna (MTF) of deep middle neritic to the upper bathyal environment is interpreted.

Keywords: Foraminifera, Campanian, Paleogene, Paleogeography, Egypt, Iraq, Iran, Southern Tethys


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